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A landscape is a heterogeneous area composed of several ecosystems. The ecosystems making up a landscape generally form a mosaic of visually distinctive patches. These patches are called landscape elements. Landscape ecology is the study of landscape structure and processes.

Landscape structure includes the size, shape, composition, number, and position of different ecosystems within a landscape. Most questions in landscape ecology require that ecologists quantify landscape structure. Until recently, however, geometry, which means "earth measurement," could offer only rough approximations of complex landscape structure. Today, an area of mathematics called fractal geometry can be used to quantify the structure of complex natural shapes. One of the findings of fractal geometry is that the length of the perimeter of complex shapes depends upon the size of the device used to measure the perimeter. One implication of this result is that organisms of different sizes may use the environment in very different ways.

Landscape structure influences processes such as the flow of energy, materials, and species between the ecosystems within a landscape. Landscape ecologists have proposed that landscape structure, especially the size, number, and isolation of habitat patches, can influence the movement of organisms between potentially suitable habitats. The group of subpopulations living on such habitat patches make up a metapopulation. Studies of the movements of small mammals in a prairie landscape show that a smaller proportion of individuals moves in more fragmented landscape but that the individuals that do move will move farther. The local population density of the Glanville fritillary butterfly, Melitaea cinxia, is lower on larger and on isolated habitat patches. Small populations of this butterfly and desert bighorn sheep are more vulnerable to local extinction. Habitat corridors have been shown to increase rates of movement among isolated habitat patches. The source of water for lakes in a Wisconsin lake district is determined by their positions in the landscape, which in turn determine their hydrologic and chemical responses to drought.

Landscapes are structured and change in response to geological processes, climate, activities of organisms, and fire. Geological features produced by processes such as volcanism, sedimentation, and erosion interact with climate to provide a primary source of landscape structure. In the Sonoran Desert, plant distributions map clearly onto soils of different ages and form a vegetative mosaic that closely matches soil mosaics. This mosaic will gradually shift as geological processes and climate gradually change the soil mosaic. While geological processes and climate set the basic template for landscape structure, the activities of organisms, from plants to elephants, can be an additional source of landscape structure and change. Economically motivated human activity changes the structure of landscapes all over the globe. Beavers can quickly change landscape structure and processes over large regions. Fire contributes to the structure of landscapes ranging from tropical savanna to boreal forest. However, fire plays a particularly prominent role in regions with a Mediterranean climate.

Because human activity has often altered landscape structure and processes in undesirable ways, there is growing pressure and interest in landscape restoration. Some of the most ambitious current restoration efforts focus on the restoration of riverine landscapes. Rivers and their floodplains form a complex, highly dynamic landscape that includes river, riparian forest, marsh, oxbow lake, and wet meadow ecosystems. Over the past century, water management by building dams, channelizing rivers, constructing flood levees, and diverting water for irrigation has cut the historic connections between most rivers and their floodplains. The restoration of the Kissimmee River landscape is also one of the largest ecological experiments ever conducted and will be a significant test of the predictive ability of ecological theory.








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